Simon, R., Matt, F., Santillan, V., Tschapka, M., Tuttle, M. and Halfwerk, W., An ultrasound absorbing inflorescence zone enhances echo-acoustic contrast of bat-pollinated cactus flowers. Journal of Experimental Biology accepted Feb 2023


Simon, R.*, Varkevisser, J. M.*, E. Mendoza, M., Hochradel, K., Elsinga, R., Wiersma, P., Middelburg, E. Zoeter, E., Scharff, C., Riebel, K. and Halfwerk, W.H. RoboFinch: an audio-visually synchronised singing robot to study vocal imitation and multimodal communication in songbirds. Methods in Ecology and Evolution accepted Dec 2022


Varkevisser, J. M., E. Mendoza, M., Simon, R., Manet, M., W. H. Halfwerk, C. Scharff and Riebel, K. (2022). Multimodality during live tutoring is relevant for vocal learning in zebra finches. Animal Behaviour |
Rickert, D., Simon, R., von Fersen, L., Baumgartner, K., Bertsch, T., Kirschbaum, C., & Erhard, M. (2022). Saliva and Blood Cortisol Measurement in Bottlenose Dolphins (Tursiops truncatus): Methodology, Application, and Limitations. Animals, 12(1), 22. |
Simon, R.*, Bakunowski, K.*, Reyes-Vasques, A. E., Tschapka, M., Knörnschild, M., Steckel, J., & Stowell, D. (2021). Acoustic traits of bat-pollinated flowers compared to flowers of other pollination syndromes and their echo-based classification using convolutional neural networks. PLoS Computational Biology, 17(12), e1009706. |
Verreycken, E., Simon, R., Quirk-Royal, B., Daems, W., Barber, J., & Steckel, J. (2021). Bio-acoustic tracking and localization using heterogeneous, scalable microphone arrays. Communications Biology, 4(1), 1-11. |
Varkevisser, J. M.*, R. Simon*, E. Mendoza, M. How, I. van Hijlkema, R. Jin, Q. Liang, C. Scharff, W. H. Halfwerk, and K. Riebel. (2021). Adding colour-realistic video images to audio playbacks increases stimulus engagement but does not enhance vocal learning in zebra finches. Animal Cognition:1-26. |
Hüttner, T.; Dollhaeupl, S.; Simon, R.; Baumgartner, K.; von Fersen, L. (2021). Activity Budget Comparisons Using Long-Term Observations of a Group of Bottlenose Dolphins (Tursiops truncatus) under Human Care: Implications for Animal Welfare. Animals, 11, 2107. |
Velilla, E., Polajnar, J., Virant-Doberlet, M., Commandeur, D., Simon, R., Cornelissen, J.H.C., Ellers, J. and Halfwerk, W. (2020) Variation in plant leaf traits affects transmission and detectability of herbivore vibrational cues. Ecology and Evolution, 10(21), 12277-12289. |
Velilla, E., Muñoz, M., Quiroga, N., Symes, L., ter Hofstede, H.M., Page, R.A., Simon, R., Ellers, J. and Halfwerk, W., (2020) Gone with the wind: Is signal timing in a neotropical katydid an adaptive response to variation in wind-induced vibratory noise? Behavioral Ecology and Sociobiology, 74, pp.1-11. |
Hernández-Montero, J.R., Reusch, C., Simon, R., Schöner, C.R. and Kerth, G., (2020) Free-ranging bats combine three different cognitive processes for roost localization. Oecologia, pp.1-10 |
Simon, R., Rupitsch, S., Baumann, M., Wu, H., Peremans, H. and Steckel, J., (2020) Bioinspired sonar reflectors as guiding beacons for autonomous navigation. Proceedings of the National Academy of Sciences. | DOI: 10.1073/pnas.1909890117
Geipel, I., Steckel, J., Tschapka, M., Vandererelst, D., Schnitzler, H. U., Kalko, E.K.V., Peremans, H.**, and Simon, R.**, (2019) Bats Actively Use Leaves as Specular Reflectors to Detect Acoustically Camouflaged Prey, Current Biology, published online August 01 |


Halfwerk, W., Varkevisser, J., Simon, R., Mendoza, E., Scharff, C., & Riebel, K. (2019) Toward Testing for Multimodal Perception of Mating Signals. Front. Ecol. Evol., 17 April 2019 |


Kroh, P., Simon, R., Rupitsch, S. J., Classification of Sonar Targets in Air: A Neural Network Approach. Sensors, 2019, 19. Jg., Nr. 5, S. 1176. |


Weber, N., Nagy, M., Hochradel, K., Mages, J., Naucke, A., Schneider, A., Stiller, F., Behr, O. & Simon, R. (2018). Akustische Erfassung der Fledermausaktivität an Windenergieanlagen. In: Bestimmung des Kollisionsrisikos von Fledermäusen an Onshore- Windenergieanlagen in der Planungspraxis – Endbericht des Forschungsvorhabens gefördert durch das Bundesministerium für Wirtschaft und Energie. Förderkennzeichen 0327638E). O. Behr et al. Erlangen / Freiburg / Ettiswil.

Behr O, Brinkmann R, Hochradel K, Mages J, Korner-Nievergelt F, Niermann I, Reich M, Simon R, Weber N, Nagy M (2017) Mitigating Bat Mortality with Turbine-Specific Curtailment Algorithms: A Model Based Approach. In Wind Energy and Wildlife Interactions (pp. 135-160). Springer International Publishing. |


Schöner, M. G., Simon, R., & Schöner, C. R. (2016). Acoustic communication in plant–animal interactions. Current opinion in plant biology, 32, 88-95. |


Übernickel, K., Simon, R., Kalko, E. K., & Tschapka, M. (2016). Sensory challenges for trawling bats: Finding transient prey on water surfaces. The Journal of the Acoustical Society of America, 139(4), 1914-1922. |


Behr, O., Brinkmann, R., Korner-Nievergelt, F., Nagy, M., Niermann, I., Reich, M.,  & Simon, R., (Hrsg., 2016) Reduktion des Kollisionsrisikos von Fledermäusen an Onshore Windenergieanlagen (RENEBAT II) – Hannover : Repositorium der Leibniz Universität Hannover, 2016 (Umwelt und Raum ; 7), 369 S. DOI:


Schöner*, M. G., Schöner*, C. R., Simon*, R. , Grafe, T. U., Puechmaille, S. J., Ji, L. L., Kerth, G. (2015). Bats Are Acoustically Attracted to Mutualistic Carnivorous Plants. Current Biology 25 (14), p. 1911–1916. |


Simon, R., Knörnschild, M., Tschapka, M., Schneider, A., Passauer, N., Kalko, E. K. and von Helversen, O. (2014). Biosonar resolving power: Echo-acoustic perception of surface structures in the submillimeter range. Frontiers in Physiology 5.


Simon, R., Holderied, M. W., Koch, C. U. and von Helversen, O. (2011) Floral acoustics: conspicuous echoes of a dish-shaped leaf attract bat pollinators. Science 333 (6042), p.631 – 633 | DOI: 10.1126/science.1204210 


Kelm DH; Simon R, Kuhlow R, Voigt CC, Ristow M, (2011) High activity enables life on a high sugar diet in nectar-feeding bats, Proceedings of the Royal Society B: Biological Sciences, 278(1724):3490-3496.


Simon, R., Holderied, M. W. and von Helversen, O., (2006). Size discrimination of hollow hemispheres by echolocation in a nectar feeding bat. Journal of Experimental Biology 209:3599-3609.



Simon, R., Dreissen, A., Leroy, H., Berg, M., & Halfwerk, W. (2022). Stealth coating enables for larger body size in a predator-prey arms race. Authorea Preprints. |

Simon, R., Matt, F., Santillan, V., Tschapka, M., Tuttle, M. and Halfwerk, W., (2019). An ultrasound absorbing inflorescence zone enhances echo-acoustic contrast of bat-pollinated cactus flowers. bioRxiv.


Simon, R., Varkevisser, J., Mendoza, E., Hochradel, K., Scharff, C., Riebel, K., & Halfwerk, W. (2019). Development and application of a robotic zebra finch (RoboFinch) to study multimodal cues in vocal communication (No. e28004v1). PeerJ Preprints.



Selected Conference Contributions


Kroh, P. K., Simon, R., & Rupitsch, S. J. (2018). Classification of Sonar Targets in Air—A Neural Network Approach. In Multidisciplinary Digital Publishing Institute Proceedings (Vol. 2, No. 13, p. 929).


Simon, R., Matt F., Holderied, M. W., Tschapka. M. (2013) Echo-acoustic signaling in bat-pollinated flowers. Acta Chiropterologica 15(1):267-269.


Behr O, Hochradel K, Mages J, Nagy M,  Korner-Nievergelt F, Niermann I, Simon R, Weber N and Brinkmann R (2013) Reducing bat fatalities at wind turbines in central Europe – How efficient are bat-friendly operation algorithms in a field-based experiment?  Conference on Wind Power and Environmental Impacts Stockholm 5-7 February


Simon R & von Helversen O (2010) Echo acoustic properties of bat-pollinated flowers and their recognition by glossophagine bats. 15th International Bat Research Conference (IBRC), Prague, Czech Republic


Simon R & von Helversen O (2007) Echo acoustic adaptations of chiropterophilous flowers. 14th International Bat Research Conference (IBRC), Merída, Mexico


Simon R & von Helversen O (2004) Size discrimination of hollow hemispheres byecholocation in the nectar feeding bat Glossophaga soricina. 13th International Bat Research Conference (IBRC), Mikołajki, Poland


von Helversen D, Simon R, von Helversen O (2003) Discrimination of rotary hollow forms by echolocation in the nectar-feeding bat Glossophaga soricina. 29th Neurobiology Conference Göttingen, Germany